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In the tradition of these meetings, Extremophiles2018, in the volcanic island of Ischia, aims to show the updated state-of-the-art of research on basic and applied aspects of life in extreme environments and to stimulate high quality research, inspiring those already working in the field and young scientists approaching extremophiles. The Conference will include sessions on many aspects of research related to extremophiles, including origin of life, ecology, astrobiology, molecular biology, physiology, and biotechnology.