ИСТИНА

The aim of our study was to research morphology and phylogenetic placement of Uzbekistanian local endemic species. Oenanthe is not typical for Middle Asia. O. heterococca was collected only once in 1927 and have never been found later. The only known herbarium specimen of Oenanthe heterococca is stored in the herbarium of Uzbekistan (TASH!). Korovin described O. heterococca and indicated heteromericarpic fruits as an important diagnostic character. We have studied fruit morphology and anatomy in O. heterococca and O. silaifolia and found both species very similar in general morphology. They have large calyx teeth, and “sepal” mericarp ribs are often larger than “petal” ribs resulting in slight heteromericarpy. Every umbel has homomericarpic and slightly heteromericarpic fruits. Analysis of ITS nrDNA for 16 species Oenanthe from GENBANK with critical specimen and other species from tribe Oenantheae showed, that O. heterococca forms with all Oenanthe species a compact separate well supported clade. Identity of Oenanthe heterococca and O. silaifolia ITS sequences is in accordance with the morphological and carpological data. Thus, O. heterococca should be synonymized with O. silaifolia and excluded from rare species of Uzbekistan. The appearance and disappearance of O. silaifolia in Uzbekistan should be regarded as the result of random drift. Two species of Autumnalia [1] form a highly supported subclade in Ferula clade. Sphaerosciadium denaense was described by Pimenov & Kljuykov [2]. Earlier Sphaerosciadium was treated as Physospermum [3]. Sphaerosciadium is situated in Komarovia-clade, the affinity with Physospermum has not been confirmed