Аннотация:The cytological and embryological description of macrosporogenesis and of formation of the embryo sacs in the lady"s mantle is presented by the case of Alchemilla filicaulis Buser. The multicellular archesporium is transformed into the reproductive complex of cells. In different flowers of the same inflorescence one can observe the septemheterotypic aneuspory, the generative apospory, and, seldom, euspory. The embryo sacs develop according to the Hieracium-type, the Taraxacum-type, and, seldom, Polygonum-type. Cases of transition from the sporophyte to the gametophyte with possible change of genotype are distinquished in scheme of processes of the archesporium transformatuon in the of macrosporogenesis in amphimicts and apomicts.